[1] Pedag Univ Krakow, Inst Biol, PL-31054 Krakow, Poland
[2] Polish Acad Sci, Inst Systemat & Evolut Anim, PL-31016 Krakow, Poland
[3] Rzeszow Univ, Dept Environm Biol, PL-35601 Rzeszow, Poland
[4] Natl Museums Scotland, Natl Museums Collect Ctr, Dept Nat Sci, Edinburgh EH5 1JA, Midlothian, Scotland