[1] South China Agr Univ, Minist Agr Peoples Republ China, Key Lab Agroenvironm Trop, Guangzhou 510642, Guangdong, Peoples R China
[2] Guangdong Acad Agr Sci, Tea Res Ctr, Drinkable Plants Inst, Guangzhou 510640, Guangdong, Peoples R China
[3] Senckenberg Res Inst, D-60325 Frankfurt, Germany
[4] Hubei Univ, Coll Life Sci, Ctr Anim Behav & Evolut, Wuhan 430062, Hubei, Peoples R China