[1] Univ Birmingham, Birmingham Med Sch, Birmingham, England
[2] Imperial Coll London, Imperial Coll, Sch Med, London, England
[3] Univ Edinburgh, Edinburgh Med Sch, Edinburgh, Scotland
[4] Univ Cambridge, Wellcome MRC Inst Metab Sci, Cambridge, England
[5] Cambridge Univ Hosp, NIHR Cambridge Biomed Res Ctr, Cambridge, England
[6] Univ Sussex, Brighton & Sussex Med Sch, Brighton, England