[1] Michigan Technol Univ, Coll Forest Resources & Environm Sci, Houghton, MI 49931 USA
[2] Univ Nebraska Omaha, Dept Biol, Omaha, NE USA
[3] Michigan Technol Univ, Michigan Tech Res Inst, Ann Arbor, MI USA
[4] Colorado State Univ, Dept Ecosyst Sci & Sustainabil, Ft Collins, CO USA
[5] United States Dept Agr USDA Forest Serv, Climate Fire & Carbon Cycle Sci Unit, Northern Res Stn, Houghton, MI USA