[1] Mannheim Univ Appl Sci, Inst Math Biol, Fac Comp Sci, D-68163 Mannheim, Germany
[2] Univ Strasbourg, Theoret Bioinformat, ICube, CNRS, 300 Blvd Sebastien Brant, F-67400 Illkirch Graffenstaden, France
[3] Univ Lorraine, CNRS, LORIA Orpailleur, Vandoeuvre les Nancy, France
[4] NRIA, Vandoeuvre les Nancy, France
[5] Radboud Univ Nijmegen, Dept Math, Nijmegen, Netherlands