[1] Guangxi Acad Sci, Guangxi Acad Marine Sci, Guangxi Key Lab Marine Environm Sci, Nanning 530007, Peoples R China
[2] Beibu Gulf Marine Ind Res Inst, Fangchenggang 538000, Peoples R China
[3] Guangxi Univ, Sch Resources Environm & Mat, Nanning 530004, Peoples R China
[4] Xiamen Univ, Sch Life Sci, Xiamen 361102, Peoples R China
[5] Chinese Acad Sci, Inst Oceanol, Lab Marine Organism Taxon & Phylogeny, Qingdao Key Lab Marine Biodivers & Conservat, Qingdao 266071, Peoples R China