[1] Shenyang Normal Univ, Coll Paleontol, Shenyang, Peoples R China
[2] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing, Peoples R China
[3] Minist Nat Resources, Key Lab Evolut Life NE Asia, Shenyang, Peoples R China
[4] Hami Museum, Hami, Peoples R China
[5] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing, Peoples R China