[1] Chinese Acad Sci, Inst Zool, Beijing 100101, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 100101, Peoples R China
[3] ARS, Systemat Entomol Lab, Natl Museum Nat Hist, USDA, C-O Smithsonian Inst, Washington, DC USA
[4] Shenzhen Polytech Univ, Plant Protect Res Ctr, Shenzhen 518055, Guangdong, Peoples R China
[5] Guangdong Acad Sci, Inst Zool, Guangdong Key Lab Anim Conservat & Resource Utiliz, Guangdong Publ Lab Wild Anim Conservat & Utilizat, Guangzhou 510260, Guangdong, Peoples R China