[1] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing, Peoples R China
[2] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing, Peoples R China
[3] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing 100044, Peoples R China
[4] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing 100049, Peoples R China