[1] Max Planck Inst Anim Behav, Dept Collect Behav, D-78464 Constance, Germany
[2] Univ Konstanz, Ctr Adv Study Collect Behav, D-78464 Constance, Germany
[3] Univ Konstanz, Dept Biol, D-78464 Constance, Germany
[4] Univ Strathclyde, Dept Naval Architecture Ocean & Marine Engn, Glasgow G4 0LZ, Scotland