[1] Univ Laval, Ctr Northern Studies CEN, Dept Geog, Quebec City, PQ, Canada
[2] Univ Laval, Takuvik Int Res Lab, Quebec City, PQ, Canada
[3] Univ Barcelona, Dept Geog, Catalonia, Spain
[4] Univ Laval, Ctr Northern Studies CEN, Takuvik Int Res Lab, Dept Biol, Quebec City, PQ, Canada
[5] Univ Laval, Inst Integrat Biol & Syst IBIS, Quebec City, PQ, Canada