[1] Univ Fed Goias, Programa Pos Grad Biodiversidade Anim, Goiania, GO, Brazil
[2] Univ Fed Espirito Santo, Dept Oceanog & Ecol, Vitoria, ES, Brazil
[3] NOAA Southwest Fisheries Sci Ctr, La Jolla, CA 92037 USA
[4] Univ Fed Espirito Santo, Dept Oceanog & Ecol, Grp Ecol Bent, Vitoria, ES, Brazil