[1] Huazhong Agr Univ, Coll Hort & Forestry Sci, Natl Key Lab Germplasm Innovat & Utilizat Crops, Wuhan 430070, Peoples R China
[2] Hubei Hongshan Lab, Wuhan 430070, Peoples R China
[3] Univ Birmingham, Sch Biosci, Coll Life & Environm Sci, Birmingham B15 2TT, England