[1] Polish Acad Sci, Inst Nat Conservat, PL-31120 Krakow, Poland
[2] Univ Huelva, Fac Ciencias Expt, Dept Ciencias Integradas, Ctr Estudios Avanzados Fis Matemat & Computac, Huelva 21071, Spain
[3] CSIC, Estn Biol Donana, Seville 41092, Spain
[4] Univ Leon, Fac Vet, Dept Sanidad Anim, Leon 24071, Spain
[5] Univ Oviedo, Biodivers Res Inst, Consejo Super Invest Cient, Principal Asturias, Mieres 33600, Spain
[6] Univ Alicante, Dept Ecol, Alicante 03690, Spain