[1] Southern Illinois Univ, Sch Biol Sci, Life Sci 2, Carbondale, IL USA
[2] Lewis & Clark Coll, Dept Biol, Portland, OR 97219 USA
[3] Yale Univ, Yale Sch Environm, New Haven, CT USA
[4] Pontificia Univ Catolica Ecuador, Escuela Ciencias Biol, Lab Ecol Plantas, Quito, Ecuador
[5] Univ Aberdeen, Sch Biol Sci, Aberdeen, Scotland
[6] Nat Hist Museum, Dept Bot, London, England
[7] Smithsonian Trop Res Inst, Panama City, Panama