[1] Amer Museum Nat Hist, Div Invertebrate Zool, Res Associate, New York, NY 10024 USA
[2] Amer Museum Nat Hist, Div Invertebrate Zool, Res Associate, Nameshi 2-33, Nagasaki 8528061, Japan
[3] Univ Hyogo, Museum Nat & Human Act, Inst Nat & Environm Sci, Yayoigaoka 6 Chome, Sanda, Hyogo 6691546, Japan
[4] Natl Museum Nat Sci, Dept Biol, 1 Guancian Rd, Taichung, Taiwan