[1] Univ Distrital Francisco Jose de Caldas, Herbario Forestal UDBC, Carrera 5 Este 15-82, Bogota, Colombia
[2] Univ Gothenburg, Dept Biol & Environm Sci, POB 463, SE-40530 Gothenburg, Sweden
[3] Univ Montpellier, CNRS, CIRAD, INRA,AMAP Lab,IRD, F-34398 Montpellier, France
[4] IRD, Herbier Guyane, AMAP Lab, BP 90165, F-97323 Cayenne, French Guiana, France