[1] Agr & Forestry Univ, Fac Agr, Chitwan, Nepal
[2] Univ Fed Rio Grande do Sul, Fac Agron, Dept Plant Hlth, Porto Alegre, Brazil
[3] Univ Fed Rio Grande do Sul, Dept Anim Sci, Anim Sci Res Program, Porto Alegre, Brazil
[4] Univ Bristol, Sch Biol Sci, 24 Tyndall Ave, Bristol BS8 1TQ, England