[1] SANParks, Sci Serv, POB 110040,Hadison Pk, ZA-8306 Kimberley, South Africa
[2] UNISA, Appl Behav Ecol & Ecosyst Res Unit, P-Bag X6,FL, ZA-1710 Pretoria, South Africa
[3] Directorate Land Use & Soil Management, Dept Agr Land Reform & Rural Dev, Pretoria 0001, South Africa
[4] Nelson Mandela Univ, Fac Sci, Nat Resource Sci & Management Cluster, P-Bag X6531, ZA-6530 George, South Africa