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Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
Shenyang Normal Univ, Liaoning Key Lab Evolut & Biodivers, Shenyang 110034, Peoples R China Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China