[1] Univ Nacl Mayor San Marcos, Fac Ciencias Biol, Unidad Posgrad, Lima, Peru
[2] Univ Nacl Mayor San Marcos, Fac Ciencias Biol, Inst Invest Ciencias Biol Antonio Raimondi, Lima, Peru
[3] Museo Hist Nat, Dept Dicotiledoneas, Lab Florist, Ave Arenales 1256, Lima, Peru
[4] Herbario AMO, Montanas Calizas 490, Miguel Hidalgo 11000, Cdmx, Mexico