DYNAMIC STUDY OF A RATIO-DEPENDENT PREDATOR-PREY MODEL WITH STRONG ALLEE EFFECT AND NONLINEAR HARVESTING

The harvesting of species occurs in terrestrial and aquatic habitats across the world. It not only causes alteration in the population structure of the species subjected to harvesting but also of the species in interaction with the harvested species. The present work investigates the effect of nonlinear prey harvesting on the dynamics of a ratio-dependent predator-prey system with a strong Allee effect in prey population. It is found that the system exhibits a rich spectrum of dynamics including saddle-node bifurcation, Hopf bifurcation and homoclinic bifurcation with respect to the parameters that shape the nonlinear harvesting rate, namely, the maximum harvesting rate and a half-saturation constant that represents the prey density at which half of the maximum harvesting rate is reached. It is found that the basin of attraction of the stable coexistence state shrinks as the harvesting rate increases and if the harvesting rate is above a threshold value at which saddle-node bifurcation occurs, the stable coexistence of predator and prey populations is not possible for any initial start. It is also found that the harvesting policies in which the harvesting rate increases less rapidly at low prey population size are more favorable for the stable coexistence of species. The presence of Allee effect in the prey population is found to increase the chances of extinction of both species by reducing the threshold value of the harvesting rate at which the unconditional extinction occurs. Numerical simulations are carried out to support the analytical findings.