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Chinese Acad Agr Sci, Inst Grassland Res, Minist Agr & Rural Affairs, Key Lab Biohazard Monitoring Green Prevent & Contr, Hohhot 010010, Peoples R China Chinese Acad Agr Sci, Inst Grassland Res, Minist Agr & Rural Affairs, Key Lab Biohazard Monitoring Green Prevent & Contr, Hohhot 010010, Peoples R China