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ONIRIS Nantes Atlantic Natl Coll Vet Med Food Sci, Dept Reprod Pathol, Site Chantrerie,BP 40706, F-44307 Nantes, France ONIRIS Nantes Atlantic Natl Coll Vet Med Food Sci, Dept Reprod Pathol, Site Chantrerie,BP 40706, F-44307 Nantes, France