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- [9] Morphological Variation in the Cryptic Species Desmognathus quadramaculatus (Black-bellied Salamander) and Desmognathus folkertsi (Dwarf Black-bellied Salamander)JOURNAL OF HERPETOLOGY, 2013, 47 (03) : 471 - 479Camp, Carlos D.Seymour, Zackary L.Wooten, Jessica A.