[1] Univ Toronto, Dept Ecol & Evolutionary Biol, 25 Willcocks St, Toronto, ON M5S 3B2, Canada
[2] Corp Colombia Hechos, Bogota, Colombia
[3] Univ Nacl Colombia, Inst Ciencias Nat, Lab Anfibios, Apartado Aereo 7495, Bogota, Colombia
[4] Univ Sao Paulo, Dept Zool, Inst Biociencias, Caixa Postal 11-461, BR-05422970 Sao Paulo, Brazil