[1] Mae Fah Luang Univ, Ctr Excellence Fungal Res, Chiang Rai 57100, Thailand
[2] Mae Fah Luang Univ, Sch Sci, Chiang Rai 57100, Thailand
[3] Minist Forestry & Agr, Ecol Div, Biotechnol & Ecol Inst, POB 811, Viangchan, Laos
[4] Chiang Mai Univ, Fac Sci, Dept Biol, Chiang Mai 50200, Thailand
[5] Univ Calif Berkeley, UC Herbarium, Berkeley, CA 94720 USA