[1] Chinese Acad Sci, Key Lab Vertebrate Evolut & Human Origins, Beijing 100044, Peoples R China
[2] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Beijing 100044, Peoples R China
[3] Ctr Excellence Life & Paleoenvironm, Beijing 100044, Peoples R China
[4] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[5] Amer Museum Nat Hist, Div Paleontol, New York, NY 10024 USA