[1] Univ Tunis El Manar, Fac Sci Tunis, Tunis 2092, Tunisia
[2] Ctr Biotechnol Borj Cedria CBBC, Lab Legumes, BP 901, Hammam Lif 2050, Tunisia
[3] Inst Natl Grandes Cultures, BP 120, Bou Salem 8170, Tunisia
[4] Ctr Biotechnol Borj Cedria CBBC, Lab Extremophile Plants, BP 901, Hammam Lif 2050, Tunisia
[5] Univ Jendouba, Higher Inst Biotechnol Beja, BP 382, Beja 9000, Tunisia
[6] Ctr Biotechnol Borj Cedria CBBC, Lab Olive Biotechnol, BP 901, Hammam Lif 2050, Tunisia