[1] Inst Marine & Antarctic Studies, Hobart, Tas, Australia
[2] Univ Tasmania, Ctr Marine Socioecol, Hobart, Tas, Australia
[3] Univ Edinburgh, Sch Geosci, Edinburgh, Midlothian, Scotland
[4] Charles Darwin Fdn, Puerto Ayora, Ecuador
[5] Charles Darwin Fdn, Charles Darwin Res Stn, Puerto Ayora, Ecuador
[6] Oceanogami, Barcelona, Spain
[7] Univ Glasgow, Inst Biodivers Anim Hlth & Comparat Med, Glasgow, Lanark, Scotland