[1] Pacific NW Natl Lab, Atmospher Sci & Global Change Div, Richland, WA 99352 USA
[2] CALTECH, Dept Comp & Math Sci, Pasadena, CA 91125 USA
[3] Carnegie Inst Sci, Dept Global Ecol, Stanford, CA 94305 USA
[4] Univ Lancaster, Lancaster Environm Ctr, Lancaster LA1 4YQ, England