[1] Univ Calif Santa Cruz, Inst Marine Sci, Santa Cruz, CA 95064 USA
[2] Rutgers State Univ, Inst Marine & Coastal Sci, New Brunswick, NJ 08903 USA
[3] Univ Calif Santa Cruz, Ocean Sci Dept, Santa Cruz, CA 95064 USA
[4] Univ Maine, Sch Marine Sci, Orono, ME USA
[5] Monterey Bay Aquarium Res Inst, Moss Landing, CA USA