[1] German Ctr Integrat Biodivers Res iDiv, Deutsch Pl 5e, D-04103 Leipzig, Germany
[2] Univ Leipzig, Inst Biol, Systemat Bot & Funct Biodivers, Johannisallee 21-23, D-04103 Leipzig, Germany
[3] Max Planck Inst Biogeochem, Hans Knoll Str 10, D-07745 Jena, Germany
[4] Univ Bayreuth, Bayreuth Ctr Ecol & Environm Res BayCEER, Dept Plant Ecol, Univ Str 31, D-95447 Bayreuth, Germany
[5] Smithsonian Trop Res Inst, Apartado 0843-03092, Balboa, Ancon, Panama