[1] Pacific NW Natl Lab, Marine Sci Lab, Sequim, WA 98382 USA
[2] Natl Sun Yat Sen Univ, Dept Marine Environm & Engn, Kaohsiung 80424, Taiwan
[3] Natl Sun Yat Sen Univ, Asia Pacific Ocean Res Ctr, Kuroshio Res Grp, Kaohsiung 80424, Taiwan
[4] Texas A&M Univ Galveston, Dept Marine Sci, Galveston, TX 77554 USA
[5] Texas A&M Univ, Dept Oceanog, College Stn, TX 77843 USA
[6] Acad Sinica, Inst Earth Sci, Taipei 11529, Taiwan
[7] Natl Sun Yat Sen Univ, Dept Oceanog, Kaohsiung 80424, Taiwan