[1] MPG Ranch, Missoula, MT 59801 USA
[2] Univ Montana, Dept Ecosyst & Conservat Sci, Missoula, MT 59812 USA
[3] Univ Chicago, Grad Program Biophys Sci, Chicago, IL 60637 USA
[4] Argonne Natl Lab, Inst Genom & Syst Biol, Lemont, IL 60439 USA
[5] Univ Copenhagen, Dept Biol, DK-2100 Copenhagen, Denmark