[1] Univ British Columbia, St Pauls Hosp, McDonald Res Lab, Vancouver, BC V6Z 1Y6, Canada
[2] Washington Univ, Sch Med, Dept Med, St Louis, MO 63110 USA
[3] Washington Univ, Sch Med, Dept Cell Biol & Physiol, St Louis, MO 63110 USA
[4] Barnes Jewish Hosp, St Louis, MO 63110 USA