[1] Seoul Natl Univ, Sch Biol Sci, Seoul 08826, South Korea
[2] Acad Sinica, Biodivers Resecurh Ctr, Taipei 115, Taiwan
[3] MNHN, Dept Systemat & Evolut, 43 Rue Cuvier,CP 26, F-75005 Paris, France
[4] MNHN, Direct Collect Invertebres, CP 53,61 Rue Buffon, F-75231 Paris 05, France
[5] Bar Ilan Univ, Mina & Everani Goodman Fac Life Sci, IL-5290002 Ramat Gan, Israel