[1] Natl Taiwan Univ, Inst Oceanog, Taipei 10764, Taiwan
[2] Natl Taiwan Normal Univ, Dept Life Sci, Taipei, Taiwan
[3] Chinese Naval Acad, Dept Oceanog, Kaohsiung, Taiwan
[4] Woods Hole Oceanog Inst, Woods Hole, MA 02543 USA
[5] Univ Washington, Appl Phys Lab, Seattle, WA 98105 USA
[6] Univ Calif San Diego, Scripps Inst Oceanog, La Jolla, CA 92093 USA
[7] Natl Taiwan Univ, Inst Oceanog, Taipei 10764, Taiwan