[1] South African Natl Biodivers Inst, Compton Herbarium, Private Bag X7, ZA-7735 Cape Town, South Africa
[2] Univ Western Cape, Dept Biodivers & Conservat Biol, Private Bag x17, ZA-7535 Cape Town, South Africa
[3] Univ Johannesburg, Dept Bot & Plant Biotechnol, POB 524, ZA-2006 Johannesburg, South Africa
[4] Univ KwaZulu Natal, Sch Biol & Conservat Sci, Res Ctr Plant Growth & Dev, Private Bag X01, ZA-3209 Pietermaritzburg, Scottsville, South Africa