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Northwest Univ, Coll Life Sci, Minist Educ,Shaanxi Key Lab Anim Conservat, Key Lab Resource Biol & Biotechnol Western China, Xian, Shaanxi, Peoples R China Northwest Univ, Coll Life Sci, Minist Educ,Shaanxi Key Lab Anim Conservat, Key Lab Resource Biol & Biotechnol Western China, Xian, Shaanxi, Peoples R China