[1] Univ Sao Paulo, Dept Bot, Inst Biociencias, BR-05422970 Sao Paulo, Brazil
[2] Univ Michigan, Herbarium, Ann Arbor, MI 48109 USA
[3] Univ Michigan, Dept Ecol & Evolutionary Biol, Ann Arbor, MI 48109 USA
[4] Secretaria Meio Ambiente, Inst Bot, BR-01061970 Sao Paulo, Brazil