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Max Planck Inst Evolutionary Biol, Dept Evolutionary Ecol, D-24306 Plon, Germany
Courant Res Ctr Evolut Social Behav, Jr Res Grp Evolut Cooperat & Prosocial Behav, D-37077 Gottingen, Germany Max Planck Inst Evolutionary Biol, Emmy Noether Grp Evolutionary Dynam, D-24306 Plon, Germany