[1] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Evolutionary Systemat Vertebrates, Beijing 100044, Peoples R China
[2] Univ Witwatersrand, Sch Med, Sch Anat Sci, ZA-2193 Johannesburg, South Africa
[3] Washington Univ, Dept Anthropol, St Louis, MO 63130 USA