[1] Simon Fraser Univ, Dept Biol Sci, Burnaby, BC V5A 1S6, Canada
[2] Smithsonian Inst, Tennenbaum Marine Observ Network, Edgewater, MD 21037 USA
[3] James Cook Univ, ARC Ctr Excellence Coral Reef Studies, Townsville, Qld 4811, Australia
[4] James Cook Univ, Coll Sci & Engn, Townsville, Qld 4811, Australia
[5] Univ Perpignan, PSL Univ Paris, CNRS, EPHE,UPVD,USR 3278,CRIOBE, F-66860 Perpignan, France
[6] Lab Excellence CORAIL, Perpignan, France
[7] Smithsonian Inst, Natl Museum Nat Hist, Dept Invertebrate Zool, Washington, DC 20560 USA
[8] Univ Washington, Sch Aquat & Fishery Sci, Seattle, WA 98105 USA
[9] Univ Washington, Burke Museum Nat Hist & Culture, Seattle, WA 98105 USA
[10] Univ New England, Sch Environm & Rural Sci, Funct Evolut & Anat Res Lab, Armidale, NSW 2351, Australia
[11] Univ New England, Sch Environm & Rural Sci, Palaeosci Res Ctr, Armidale, NSW 2351, Australia
[12] Smithsonian Inst, Natl Museum Nat Hist, Dept Vertebrate Zool, Washington, DC 20560 USA