[1] Univ Sao Paulo, Inst Oceanog, Dept Oceanog Biol, Lab Dinam Bent, 191 Sala 139, BR-05508120 Sao Paulo, SP, Brazil
[2] Univ Estado Rio de Janeiro, Inst Biol, Dept Zool, Lab Zool Invertebrados, Sala 516,Rua Sao Francisco Xavier 524, BR-20550900 Rio De Janeiro, RJ, Brazil
[3] Japan Agcy Marine Earth Sci & Technol JAMSTEC, 2-15 Natsushima Cho, Yokosuka, Kanagawa 2370061, Japan