[1] Natl Sun Yat Sen Univ, Inst Marine Resources, Kaohsiung 80424, Taiwan
[2] Taiwan Forestry Res Inst, Div Silviculture, Taipei, Taiwan
[3] Natl Pingtung Univ Sci & Technol, Pingtung 912, Taiwan
[4] Natl Sun Yat Sen Univ, Inst Biol, Kaohsiung, Taiwan