[1] Univ Paris Saclay, BREED, ENVA, INRAE,UVSQ, F-78350 Jouy En Josas, France
[2] Inst Pasteur, Lab Genet Dev Humain, UMR 3738, Biol Dev & Cellules Souches, 25 Rue Docteur Roux, F-75015 Paris, France
[3] Swansea Univ, Med Sch, Singleton Pk, Swansea SA2 8PP, W Glam, Wales
[4] Fac Rene Descartes, Inst Cochin, UMR CNRS 8104, INSERM U1016, 24 Rue Faubourg St Jacques, F-75014 Paris, France
[5] Obihiro Univ Agr & Vet Med, Obihiro, Hokkaido 0808555, Japan