[1] Univ New England, Nat Hist Museum, Insect Ecol Lab, Zool, Armidale, NSW 2351, Australia
[2] La Trobe Univ, Sch Life Sci, Res Ctr Future Landscapes, Dept Ecol Environm & Evolut, Bundoora, Vic, Australia
[3] Federat Univ Australia, Future Reg Res Ctr, Mt Helen, Vic, Australia