[1] Univ Gottingen, JF Blumenbach Inst Zool & Anthropol, Gottingen, Germany
[2] Univ Hong Kong, Sch Biol Sci, Hong Kong, Hong Kong, Peoples R China
[3] Tijuana River Natl Estuarine Res Reserve, Imperial Beach, CA 91932 USA
[4] Santa Barbara Museum Nat Hist, Santa Barbara, CA 93105 USA
[5] Senckenberg Meer, Dept Marine Res, D-26382 Wilhelmshaven, Germany