[1] Ecole Prat Hautes Etud, Inst Bot, Paleoenvironm & Chronoecol PALECO EPHE, F-34090 Montpellier, France
[2] Univ Montpellier 2, Inst Bot, Ctr Bioarchaeol & Ecol CNRS UMR5059, F-34090 Montpellier, France
[3] Univ Naples 2, Dept Environm Biol & Pharmaceut Sci & Technol Di, I-81100 Caserta, Italy
[4] Swiss Fed Res Inst WSL, CH-8903 Birmensdorf, Switzerland
[5] Univ Bern, Oeschger Ctr Climate Change Res, CH-3012 Bern, Switzerland
[6] Global Change Res Ctr AS CR, Vvi, Brno 60300, Czech Republic
[7] Univ Quebec, Ctr Etud Foret, Montreal, PQ H3C 3P8, Canada